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Abstract Peridotite xenoliths entrained in magmas near the Alpine fault (New Zealand) provide the first direct evidence of deformation associated with the propagation of the Australian-Pacific plate boundary through the region at ca. 25–20 Ma. Two of 11 sampled xenolith localities contain fine-grained (40–150 μm) rocks, indicating that deformation in the upper mantle was focused in highly sheared zones. To constrain the nature and conditions of deformation, we combine a flow law with a model linking recrystallized fraction to strain. Temperatures calculated from this new approach (625–970 °C) indicate that the observed deformation occurred at depths of 25–50 km. Calculated shear strains were between 1 and 100, which, given known plate offset rates (10–20 mm/yr) and an estimated interval during which deformation likely occurred (<1.8 m.y.), translate to a total shear zone width in the range 0.2–32 km. This narrow width and the position of mylonite-bearing localities amid mylonite-free sites suggest that early plate boundary deformation was distributed across at least ∼60 km but localized in multiple fault strands. Such upper mantle deformation is best described by relatively rigid, plate-like domains separated by rapidly formed, narrow mylonite zones. 

Abstract Mantle xenoliths from the Southern Alps, New Zealand, provide insight into the origin of mantle seismic anisotropy related to the Australian‐Pacific plate boundary. Most xenoliths from within 100 km lateral distance of the Alpine Fault are coarse grained, but a small number are finer grained protomylonites. The protomylonites contain connected networks of fine grains with a different crystallographic preferred orientation (CPO) to coarse porphyroclasts in the same xenolith, suggesting that protomylonites and coarse‐grained samples record different deformation kinematics. The CPOs of fine grains in protomylonites have monoclinic symmetry, with the 2‐fold rotation axis normal to a plane that contains olivine [010] and orthopyroxene [100] maxima, suggesting that the protomylonite deformation involved significant simple shear. Some coarse‐grained samples contain unconnected lenses and layers of fine grains with the same CPO as the coarse grains. Microstructures suggest that these fine grains formed by subgrain rotation recrystallization and that protomylonites may represent an up‐strain progression of this microstructure, where the connectivity of fine grains has allowed them to localize shear and develop a new Alpine Fault CPO. The samples tell us about the state of the mantle at 25 Ma, in the early history of the plate boundary. If this suite of samples is representative of the mantle beneath the Alpine Fault in the present day, then we can interpret the complex seismic anisotropy patterns in the lithospheric mantle as representative of blocks containing variably rotated older CPOs juxtaposed by narrow shear zones associated with Alpine Fault deformation.